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MALACOLOGIA
http://malacologia.fmnh.org
World’s Leading Malacological Journal

Journal Impact Factor 1.440 (2009)
Founded in 1961

EDITOR-IN-CHIEF
GEORGE M. DAVIS

Editorial Office
Malacologia
P.O. Box1222
West Falmouth, Massachusetts 02574, U.S.A.
georgedavis99@hotmail.com

Business & Subscription Office

Malacologia
P.O. Box 385
Haddonfield, New Jersey 08033, U.S.A.
malacologia@comcast.net

Copy Editor

EUGENE COAN
California Academy of Sciences
San Francisco, California, U.S.A.
gene.coan@sierraclub.org

Associate Editor

JOHN B. BURCH
University of Michigan
Ann Arbor, Michigan, U.S.A.
jbburch@umich.edu

Managing Editor

CARYL HESTERMAN
Haddonfield, New Jersey, U.S.A.
malacologia@comcast.net

Composition Editor

CLAUDIA WILKE
Giessen, Germany
claudiawilke@hotmail.com

Graphics Editor

THOMAS WILKE
Justus Liebig University
Giessen, Germany
tom.wilke@allzool.bio.uni-giessen.de

MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY

Board of Directors - U.S.A.:

President

RÜDIGER BIELER
Field Museum, Chicago, Illinois

Vice President

PAULA M. MIKKELSEN
Paleontological Research Institution, Ithaca, New York

Secretary and Treasurer

GEORGE M. DAVIS
West Falmouth, Massachusetts

Associate Member

ELLEN E. STRONG
National Museum of Natural History, Washington D.C.

Members

JOHN B. BURCH
University of Michigan, Ann Arbor

DIARMAID Ó FOIGHIL

University of Michigan, Ann Arbor

CAROLE S. HICKMAN

University of California, Berkeley

ALAN KOHN

University of Washington, Seattle

SHI-KUEI WU

University of Colorado Museum, Boulder

Emeritus Founding Consultant

J FRANCES ALLEN

Copyright © 2010 by the Institute of Malacology

ISSN: 0076-2997