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MALACOLOGIA
http://malacologia.fmnh.org

One of the World’s Leading Malacological Journals
Journal Impact Factor 1.024 (2010)

Founded in 1961

EDITOR-IN-CHIEF
GEORGE M. DAVIS

Editorial Office
Malacologia
P.O. Box 1222
West Falmouth, Massachusetts 02574, U.S.A.
e-mail

Business & Subscription Office
Malacologia
P.O. Box 385
Haddonfield, New Jersey 08033, U.S.A.
e-mail

Copy Editor
EUGENE COAN
Santa Barbara Museum of Natural History
Santa Barbara, California, U.S.A.
e-mail

Graphics Editor
THOMAS WILKE
Justus Liebig University
Giessen, Germany
e-mail

Associate Editor
JOHN B.BURCH
University of Michigan
Ann Arbor, Michigan, U.S.A.
e-mail

Managing Editor
CARYL HESTERMAN
Haddonfield, New Jersey, U.S.A.
e-mail

Composition Editor
CLAUDIA WILKE
Giessen, Germany
e-mail

MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY
Board of Directors - U.S.A.:

President
PAULA M. MIKKELSEN
Paleontological Research Institution, Ithaca, New York

Vice President
DIARMAID Ó FOIGHIL
University of Michigan, Ann Arbor

Secretary and Treasurer
GEORGE M. DAVIS
West Falmouth, Massachusetts

Members
RÜDIGER BIELER
Field Museum, Chicago, Illinois

JOHN B. BURCH
University of Michigan, Ann Arbor

CAROLE S. HICKMAN
University of California, Berkeley

ALAN KOHN
University of Washington, Seattle

ELLEN STRONG
National Museum of Natural History, Washington D.C.

Copyright © 2012 by the Institute of Malacology
ISSN: 0076-2997