Kenneth D. Angielczyk's Publications
PDF copies of all publications are available on request.
Journal Publications
Olroyd, S. L., Sidor, C. A., and Angielczyk, K. D. 2018. New materials of the enigmatic dicynodont Abajudon kaayai (Therapsida, Anomodontia) from the lower Madumabisa Mudstone Formation, middle Permian of Zambia. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1403442.
Benoit, J. Angielczyk, K. D., Miyamae, J. A., Manger, P., Fernandez, V., and Rubidge, B. 2018. Evolution of facial innervation in anomodont therapsids (Synapsida): insights from x-ray computerized microtomography. Journal of Morphology. DOI: 10.1002/jmor.20804.
Bernardi, M., Petti, F. M., Kustatscher, E., Franz, M., Hartkopf-Fröder, C., Labandeira, C. C., Wappler, T., van Konijnenburg-van Cittert, J. H. A., Peecook, B. R., and Angielczyk, K. D. 2017. Late Permian (Lopingian) terrestrial ecosystems: a global comparison with new data from the low-latitude Bletterbach Biota. Earth-Science Reviews 175: 18-43.
Angielczyk, K. D. and Kammerer, C. F. 2017. The cranial morphology, phylogenetic positoion, and biogeography of the upper Permian dicynodont Compsodon helmoedi van Hoepen (Therapsida, Anomodontia). Papers in Palaeontology 3: 513-545.
Nesbitt, S. J., Butler, R. J. Ezcurra, M. D., Barrett, P. M., Stokcer, M. R., Angielczyk, K. D., Smith, R. M. H., Sidor, C. A., Niedźwiedzki, G., Sennikov, A. G., and Charig, A. J. 2017. The earliest bird-line archosaurs and the assembly of the dinosaur body plan. Nature 544: 484-487.
Codron, J., Botha-Brink, J., Codron,D., Huttenlocker, A. K., and Angielczyk, K. D. 2017. Predator-prey interactions amongst Permo-Triassic terrestrial vertebrates as a deterministic factor influencing faunal collapse and turnover. Journal of Evolutionary Biology 30: 40-54.
Bernardi, M., Angielczyk, K.D., Mitchell, J. S., and Ruta, M. 2016. Phylogenetic stability, tree shape, and character compatibility: a case study using early tetrapods. Systematic Biology 65: 737-758.
Barbolini, N., Smith R. M. H., Tabor, N. J., Sidor, C. A., and Angielczyk K. D. 2016. Resolving the age of Madumabisa fossil vertebrates: palynological evidence from the mid-Zambezi Basin of Zambia. Palaeogeography, Palaeoclimatology, Palaeoecology 457: 117-128.
Botha-Brink, J., Codron, D., Huttenlocker, A. K., Angielczyk, K. D., and Ruta M. 2016. Breeding young as a survival strategy during Earth's greatest mass extinction. Scientific Reports 6: 24053. DOI: 10.1038/srep24053.
Polly, P. D., Stayton, C. T., Dumont, E. R., Pierce S. E., Rayfield, E. J., and Angielczyk, K. D. 2016. Combining geometric morphometrics and finite element analysis with evolutionary modeling: towards a synthesis. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1111225.
Angielczyk, K. D., Rubidge, B. S., Day, M. O., and Lin, F. 2016. A reevaluation of Brachyprosopus broomi and Chelydontops altidentalis, dicynodonts (Therapsida, Anomodontia) from the middle Permian Tapinocephalus Assemblage Zone of the Karoo Basin, South Africa. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1078342.
Viglietti, P. A., Smith, R. M. H., Angielczyk, K. D., Kammerer, C. F., Fröbisch, J., and Rubidge, B. S. 2016. The Daptocephalus Assemblage Zone (Lopingian), South Africa: a proposed biostratigraphy based on a new compilation of stratigraphic ranges. Journal of African Earth Sciences 113: 153-164.
Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. 2015. Redescription of the geikiid Pelanomodon (Therapsida, Dicynodontia), with a reconsideration of 'Propelanomodon.' Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2015.1030408.
Cisneros, J. C., Marsicano, C., Angielczyk, K. D., Smith, R. M. H, Richter, M., Fröbisch, J., Kammerer, C. F., and Sadleir, R. W. 2015. New Permian fauna from tropical Gondwana. Nature Communications 6: 8676. DOI: 10.1038/ncomms9676.
Angielczyk, K. D. and Cox, C. B. 2015. Distinctive emydopoid dicynodont (Therapsida, Anomodontia) mandibles from the Permian Ruhuhu and Usili formations (Songea Group), Ruhuhu Basin, Tanzania. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2015.1008699.
Roopnarine, P. D. and Angielczyk, K. D. 2015. Community stability and selective extinction during the Permian-Triassic mass extinction. Science 350: 90-93.
Huttenlocker, A. K, Sidor, C. A., and Angielczyk, K. D. 2015. A new eutherocephalian (Therapsida, Therocephalia) from the upper Permian Madumabisa Mudstone Formation (Luangwa Basin) of Zambia. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2015.969400.
Cox, C. B. and Angielczyk, K. D. 2015. A new endothiodont dicynodont (Therapsida, Anomodontia) from the Permian Ruhuhu Formation (Songea Group) of Tanzania and its feeding system. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2014.935388
Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. 2015. Redescription of Digalodon rubidgei, an emydopoid dicynodont (Therapsida, Anomodontia) from the Late Permian of South Africa. Fossil Record 18: 43-55.
Angielczyk, K. D., Burroughs, R. W., and Feldman, C. R. 2015. Do turtles follow the rules? Latitudinal gradients in species richness, body size, and geographic range area of the world's turtles. Journal of Experimental Zoology Part B (Molecular and Developmental Evolution) 324B:270-294.
Angielczyk, K. D., Huertas, S., Smith, R. M. H., Tabor, N. J., Sidor, C. A., Steyer, J.-S., Tsuji, L. A., and Gostling, N. J. 2014. New dicynodonts (Therapsida, Anomodontia) and updated tetrapod stratigraphy of the Permian Ruhuhu Formation (Songea Group, Ruhuhu Basin) of Southern Tanzania. Journal of Vertebrate Paleontology 34: 1408-1426.
Nesbitt, S. J., Sidor, C. A., Angielczyk, K. D., Smith, R. M. H., and Tsuji, L. A. 2014. A new archosaur from the Manda beds (Anisian, Middle Triassic) of souther Tanzania and its implications for character state optimizations at Archosauria and Pseudosuchia. Journal of Vertebrate Paleontology 34: 1357-1382.
Angielczyk, K. D. and Schmitz, L. 2014. Nocturnality in synapsids predates the origin of mammals by over 100 million years. Proceedings of the Royal Society B 281: 20141642. doi:10.1098/rspb.2014.1642.
Sidor, C. A., Angielczyk, K. D., Smith, R. M. H., Goulding, A. K., Nesbitt, S. J., Peecook, B. R., Steyer, J.-S. and Tolan, S. 2014. Tapinocephalids (Therapsida: Dinocephalia) from the Permian Madumabisa Mudstone Formation (Lower Karoo, Mid-Zambezi Basin) of southern Zambia. Journal of Vertebrate Paleontology 34: 980-986.
Castanhinha, R., Araújo, R., Júnior, L. C., Angielczyk, K. D., Martins, G. G., Martins, R. M. S., Chaouiya, C., Beckmann, F., and Wilde, F. 2013. Bringing dicynodonts back to life: paleobiology and anatomy of a new emydopoid genus from the Upper Permian of Mozambique. PLoS One 8(12): e80974. doi:10.1371/journal.pone.0080974
Peecook, B. R., Sidor, C. A., Nesbitt, S. J., Smith, R. M. H., Steyer, J. S., and Angielczyk, K. D. 2013. A new silesaurid from the Upper Ntawere Formation of Zambia (Middle Triassic) demonstrates the rapid diversification of Silesauridae (Avemetatarsalia, Dinosauriformes). Journal of Vertebrate Paleontology 33: 1127-1137.
Ruta, M. Angielczyk, K. D., Fröbisch, J., and Benton, M. J. 2013. Decoupling of morphological disparity and taxic diversity during the adaptive radiation of anomodont therapsids. Proceedings of the Royal Society Series B 280: 20131071. doi:10.1098/rspb.2013.1071.
Kammerer, C. F., Fröbisch, J., and Angielczyk, K. D. 2013. On the validity and phylogenetic position of Eubrachiosaurus browni, a kannemeyeriiform dicynodont (Anomodontia) form Triassic North America. PLoS One 8(5): e64203. doi:10.1371/journal.pone.0064203
Sidor, C. A., Vilhena, D. A., Angielczyk, K. D., Huttenlocker, A. K., Nesbitt, S. J., Peecook, B. R., Steyer, J.-S., Smith, R. M. H., and Tsuji, L. A. 2013. Provincialization of terrestrial faunas following the end-Permian mass extinction. Proceedings of the National Academy of Sciences 110: 8129-8133.
Hancox, P. J., Angielczyk, K. D., and Rubidge, B. S. 2013. Angonisaurus and Shansiodon, dicynodonts (Therapsida, Anomodontia) from subzone C of the Cynognathus Assemblage Zone (Middle Triassic) of South Africa. Journal of Vertebrate Paleontology 33: 655-676.
Angielczyk, K. D. and Feldman, C. R. 2013. Are diminutive turtles miniaturized? The ontogeny of plastron shape in emydine turtles. Biological Journal of the Linnean Society108: 727-755.
Angielczyk, K. D. and Rubidge, B. S. 2013. Skeletal morphology, phylogenetic relationships, and stratigraphic range of Eosimops newtoni Broom, 1921, a pylaecephalid dicynodont (Therapsida, Anomodontia) from the Middle Permian of South Africa. Journal of Systematic Palaeontology 11: 191-231.
Mitchell, J. S., Roopnarine, P. D., and Angielczyk, K. D. 2012. Late Cretaceous restructuring of terrestrial communities facilitated the end-Cretaceous mass extinction in North America. Proceedings of the National Academy of Sciences 109:18857-18861.
Angielczyk, K. D. and Ruta, M. 2012. The roots of amphibian morphospace: a geometric morphometric analysis of Paleozoic temnospondyls. Fieldiana: Life and Earth Sciences 5: 40-58.
Parham, J. F., Donoghue, P. C. J., Bell, C. J., Calway, T. D., Head, J. J., Holroyd, P. A., Inoue, J. G., Irmis, R. B., Joyce, W. G., Ksepka, D. T., Patané, J. S. L., Smith, N. D., Tarver, J. E., van Tuinen, M., Yang, Z., Angielczyk, K. D., Greenwood, J. M., Hipsley, C. A., Jacobs, L., Makovicky, P. J., Müller, J., Smith, K. T., Theodor, J. M., Warnock, R. C. M., and Benton, M. J. 2012. Best practices for justifying fossil calibrations. Systematic Biology 61: 346-359.
Roopnarine, P. D. and Angielczyk, K. D. 2012. The evolutionary palaeoecology of species and the tragedy of the commons. Biology Letters 8: 147-150.
Kammerer, C. F., Angielczyk, K. D., and J. Fröbisch. 2011. A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia), and its implications for dicynodont phylogeny, biogeography, and biostratigraphy. Society of Vertebrate Paleontology Memoir 11: 1-158.
Angielczyk, K. D., Feldman, C. R., and Miller, G. R. 2011. Adaptive evolution of plastron shape in emydine turtles. Evolution 65: 377-394.
Botha-Brink, J. and Angielczyk, K. D. 2010. Do extraordinarily high growth rates in Permo-Triassic dicynodonts (Therapsida, Anomodontia) explain their success before and after the end-Permian extinction? Zoological Journal of the Linnean Society 160: 341-365.
Angielczyk, K. D. and Rubidge, B. S. 2010. A new pylaecephalid dicynodont from the Tapinocephalus Assemblage Zone, Karoo Basin, Middle Permian of South Africa. Journal of Vertebrate Paleontology 30: 1396-1409.
Simon, R. V., Sidor, C. A., Angielczyk, K. D., and Smith, R. M. H. 2010. First record of a tapinocephalid (Therapsida: Dinocephalia) from the Ruhuhu Formation (Songea Group) of southern Tanzania. Journal of Vertebrate Paleontology 30: 1289-1293.
Sidor, C. A., Angielczyk, K. D., Weide, D. M., Smith, R. M. H., and Tsuji, L. A. 2010. Tetrapod fauna of the lowermost Usili Formation (Songea Group, Ruhuhu Basin) of southern Tanzania, with a new burnetiid record. Journal of Vertebrate Paleontology 30: 696-703.
Nesbitt, S. J., Sidor, C. A., Irmis, R. B., Angielczyk, K. D., Smith, R. M. H., and Tsuji, L. A. 2010. Ecologically distinct dinosaurian sister-group indicates early diversification of Ornithodira. Nature 464: 95-98.
Fröbisch, J., Angielczyk, K. D., and Sidor, C. A. 2010. The Triassic dicynodont Kombuisia (Synapsida, Anomodontia) from Antarctica, a refuge for terrestrial tetrapods during and after the end-Permian extinction. Naturwissenschaften 97: 187-196.
Weide, D. M., Sidor, C. A., Angielczyk, K. D., and Smith, R. M. H. 2009. A new record of Procynosuchus delaharpeae (Therapsida: Cynodontia) from the Upper Permian Usili Formation, Tanzania. Palaeontologia africana 44: 21-26.
Angielczyk, K. D., Sidor, C. A., Nesbitt, S. J., Smith, R. M. H., and Tsuji, L. A. 2009. Taxonomic revision and new observations on the postcranial skeleton, biogeography, and biostratigraphy of the dicynodont genus Dicynodontoides, the senior subjective synonym of Kingoria (Therapsida, Anomodontia). Journal of Vertebrate Paleontology 29: 1174-1187.
Angielczyk, K. D. and Rubidge, B. S. 2009. The Permian dicynodont Colobodectes cluveri (Therapsida, Anomodontia), with notes on its ontogeny and stratigraphic range in the Karoo Basin, South Africa. Journal of Vertebrate Paleontology 29: 1162-1173.
Pierce, S. E., Angielczyk, K. D., and Rayfield, E. J. 2009. Morphospace occupation in thalattosuchian crocodylomorphs: skull shape variation, species delineation, and temporal patterns. Palaeontology 52: 1057-1097.
Pierce, S. E., Angielczyk, K. D., and Rayfield, E. J. 2009. Shape and mechanics in thalattosuchian (Crocodylomorpha) skulls: implications for feeding behavior and niche partitioning. Journal of Anatomy 215: 555-576.
Kammerer, C. F. and Angielczyk, K. D. 2009. A proposed higher taxonomy of anomodont therapsids. Zootaxa 2018: 1-24.
Angielczyk, K. D. 2009. Dimetrodon is not a dinosaur: using tree thinking to understand the ancient relatives of mammals and their evolution. Evolution: Education and Outreach 2: 257-271. DOI: 10.1007/s12052-009-0117-4.
Angielczyk, K. D. and Sullivan, C. 2008. Diictodon feliceps (Owen, 1876), a dicynodont (Therapsida, Anomodontia) species with a Pangaean distribution. Journal of Vertebrate Paleontology 28: 788-802.
Pierce, S. E., Angielczyk, K. D., and Rayfield, E. J. 2008. Patterns of morphospace occupation and mechanical performance in extant crocodilian skulls: a geometric morphometric and finite element modeling approach. Journal of Morphology 269: 840-864.
Angielczyk, K. D. and Walsh M. L. 2008. Patterns in the evolution of nares size and secondary palate length in anomodont therapsids (Synapsida): implications for hypoxia as a cause for end-Permian terrestrial vertebrate extinctions. Journal of Paleontology 82: 528-542.
Botha, J. and Angielczyk, K. D. 2007. An integrative approach to distinguishing the Late Permian dicynodont species Oudenodon bainii and Tropidostoma microtrema (Therapsida: Anomodontia). Palaeontology 50: 1175-1209.
Roopnarine, P. D., Angielczyk, K. D., Wang, S. C., and Hertog, R. 2007. Food web models explain instability of Early Triassic terrestrial communities. Proceedings of the Royal Society Series B 274: 2077-2086.
Angielczyk, K. D. 2007. New specimens of the Tanzanian dicynodont “Cryptocynodon” parringtoni Von Huene, 1942 (Therapsida, Anomodontia), with an expanded analysis of Permian dicynodont phylogeny. Journal of Vertebrate Paleontology 27: 116-131.
Angielczyk, K. D. and Sheets, H. D. 2007. Investigation of simulated tectonic deformation in fossils using geometric morphometrics. Paleobiology 33: 125-148.
Angielczyk, K. D. and Fox, D. L. 2006. New uses for measures of the fit of phylogenetic hypotheses to the fossil record. Paleobiology 32: 147-165.
Angielczyk, K. D., Fröbisch, J., and Smith, R. M. H. 2005. On the stratigraphic range of the dicynodont taxon Emydops (Therapsida, Anomodontia) in the Karoo Basin, South Africa. Palaeontologia africana 41: 23-33.
Dayrat, B., Schander, C., and Angielczyk, K. D. 2004. Suggestions for a new species nomenclature. Taxon 53: 485-491.
Angielczyk, K. D. 2004. Phylogenetic evidence for and implications of a dual origin of propaliny in anomodont therapsids (Synapsida). Paleobiology 30: 268-296.
Angielczyk, K. D. and Kurkin, A. A. 2003. Phylogenetic analysis of Russian Permian dicynodonts (Therapsida: Anomodontia): implications for Permian biostratigraphy and Pangaean biogeography. Zoological Journal of the Linnean Society 139: 157-212.
Angielczyk, K. D. and Kurkin, A. A. 2003. Has the utility of Dicynodon for Upper Permian terrestrial biostratigraphy been overstated? Geology 31: 363-366.
Angielczyk, K. D. 2002. Character state transformations and the fit of phylogenies to the fossil record. Palaeontologia africana 38: 27-32.
Angielczyk, K. D. 2002. Redescription, phylogenetic position, and stratigraphic significance of the dicynodont genus Odontocyclops (Synapsida; Therapsida). Journal of Paleontology 76: 1047-1059.
Nesbitt, S. J. and Angielczyk, K. D. 2002. New evidence of large dicynodonts in the Upper Moenkopi Formation (Middle Triassic) of northern Arizona. PaleoBios 22: 10-17.
Angielczyk, K. D. 2002. A character-based method for measuring the fit of a cladogram to the fossil record. Systematic Biology 51: 176-191.
Angielczyk, K. D. 2001. Preliminary phylogenetic analysis and stratigraphic congruence of the dicynodont anomodonts (Synapsida: Therapsida). Palaeontologia Africana 37: 53-79.
Angielczyk, K. D. and Gingerich, P. D. 1998. New specimen of cf. Asiatosuchus (Crocodyloidea) from the Middle Eocene Drazinda Formation of the Sulaiman Range, Punjab (Pakistan). Contributions from the Museum of Paleontology, University of Michigan 30 (5): 163-189.
Edited Volumes
Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. 2014. Early Evolutionary History of the Synapsida. Springer, Dordrecht. 337pp.
Contributions to Edited Volumes
Angielczyk, K. D. and Kammerer, C. F. 2017. Non-mammalian synapsids: the deep roots of the mammalian family tree. In Zachos, F. E. and Asher, R. J. (eds.), Handbook of Zoology: Mammalia: Mammalian Evolution, Diversity, and Systematics. De Gruyter, Berlin.
Roopnarine, P. D. and Angielczyk, K. D. 2016. The stability of ecological communities as an agent of evolutionary selection. Pp. 307-333 in Eldredge, N., Pievani T., Serrelli, E., and Temkin I. (eds.), Evolutionary Theory: A Hierarchical Perspective. University of Chicago Press, Chicago.
Angielczyk, K. D. 2014. Therapsid diversity patterns and the end-Permian extinction: Introduction. Pp. 283-287 in: Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. (eds.), Early Evolutionary History of the Synapsida. Springer, Dordrecht.
Angielczyk, K. D., Steyer, J. S., Sidor, C. A., Smith, R. M. H., Whatley, R. L., and Tolan, S. 2014. Permian and Triassic dicynodont (Therapsida: Anomodontia) faunas of the Luangwa Basin, Zambia: taxonomic update and implications for dicynodont biogeography and biostratigraphy. Pp. 93-138 in: Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. (eds.), Early Evolutionary History of the Synapsida. Springer, Dordrecht.
Felice, R. N. and Angielczyk, K. D. 2014. Was Ophiacodon (Synapsida, Eupelycosauria) a swimmer? A test using vertebral dimensions. Pp. 25-51 in: Kammerer, C. F., Angielczyk, K. D., and Fröbisch, J. (eds.), Early Evolutionary History of the Synapsida. Springer, Dordrecht.
Aronowsky, A., Sanzenbacher, B., Childs, M., Peachey, A., Villanosa, K., Thompson, J., and Angielczyk, K. D. 2013. Promoting embodied learning through virtual and real world gaming experiences. Pp. 68-74 in: Williams, C. C., Ochsner, A., Dietmeier, J., and Steinkuehler, K. (eds.), Proceedings, GLS 9.0, Games + Learning + Society Conference. ETC Press, Pittsburgh.
Angielczyk, K., Aronowsky, A., Sanzenbacher, B., and Villanosa, K. 2012. Game of Bones: decisions and early feedback from a vertical slice. Pp. 63-68 in: Martin, C., Ochsner, A., and Squire, K. (eds.), Proceedings, GLS 8.0, Games + Learning + Society Conference. ETC Press, Pittsburgh.
Padian, K. and Angielczyk, K. D. 2007. “Transitional forms” versus transitional features. Pp. 197-230 in: A. J. Petto and L. Godfrey (eds.); Scientists Confront Intelligent Design and Creationism. W. W. Norton & Co., New York.
Angielczyk, K. D., Roopnarine, P. D., and Wang, S. C. 2005. Modeling the role of primary productivity disruption in end-Permian extinctions, Karoo Basin, South Africa. In: S. G. Lucas and K. E. Zeigler (eds.); The Nonmarine Permian. Bulletin of the New Mexico Museum of Natural History and Science 30: 16-23.
Angielczyk, K. D. 2002. A selective annotation of published Triassic vertebrates from the UCMP collection. In: A. B. Heckert and S. G. Lucas (eds.); Triassic Stratigraphy and Paleontology. Bulletin of the New Mexico Museum of Natural History and Science 21: 297-301.
Padian, K. and Angielczyk, K. D. 1999. Are there transitional forms in the fossil record? In: P. H. Kelley, J. R. Bryan, and T. A. Hansen (eds.); The Evolution-Creation Controversy II: Perspectives on Science, Religion, and Geological Education. Paleontological Society Papers 5: 47-82.
Encyclopedia Entries
Angielczyk, K. D. 2011. Dicynodontia. Pp. 76-79 in: 2011 McGraw-Hill Yearbook of Science & Technology. McGraw-Hill, New York.
Technical Comments
Roopnarine, P. D., Angielczyk, K. D., and Hertog, R. 2006. Comment on “Statistical independence of escalatory ecological trends in Phanerozoic marine invertebrates.” Science 314: 925d.
Book Reviews
Angielczyk, K. D. 2008. Non-Marine Permian Biostratigraphy and Biochronology, Edited by Spencer G. Lucas, Giuseppe Cassinis, and Joerg W. Schneider. Palaios. DOI: 10.2110/palo.2008.BR28.
Angielczyk, K. D. 2005. Review of Robert A. Martin, Missing Links: Evolutionary Concepts & Transitions Through Time. Reports of the National Center for Science Education 25: 42-43.
